Pyxidicula cymbalum
Pyxidicula cymbalum, diameter 90 µm

Pyxidicula cymbalum Penard, 1902

Diagnosis: Shell patelliform, transparent, usually light brown, and distinctly punctate. Aperture round, with a double rim: one curved inward and the other outward. Contractile vacuole single. Pseudopodia unknown.

Dimensions: According to Penard (1902): shell diameter 85–90 µm. My measurements over the years: 69–120 µm. Specimens from a Mongolian fen (Łuców et al., 2025): shell diameter 56–93 µm; building units 0.9–1.1 µm across; rim 7.4–8.8 µm wide (n = 21).

Ecology: Wet moss and Sphagnum, but also found in various water types, e.g. drinking‑water basins (Waterlaboratorium – Haarlem).

Geographical distribution: Probably cosmopolitan. Common, but easily overlooked.

Remarks: Although not mentioned by Penard (1902), two rims are present: an outer rim, which is always present, and an inner rim, which may be very small, absent, or difficult to detect. Both rims may form an angle of 180° or less, appearing as a V in cross‑section. The outer rim may show radiating striations or an alveolar structure (see below).

Pyxidicula cymbalum
Pyxidicula cymbalum, detail of the outer rim.
Pyxidicula cymbalum
Pyxidicula cymbalum, structure of the dorsal part of the shell.
Pyxidicula-cymbalum
Original drawing of Penard (1902) in Faune Rhizopodique du Bassin du Léman.
Pyxidicula cymbalum
The structure of the double margin. There is an inner margin and an outer rim. The inner margin is not always present or distinct. It can also be hidden by the outer rim (f2), if viewed from the dorsal side. (Drawing: Ferry Siemensma)
Pyxidicula cymbalum
Stacked images, test from Austria, 89 µm.
Pyxidicula cymbalum
Pyxidicula cymbalum, edge with distinct inner and outer rim, 92 µm
Pyxidicula cymbalum
Pyxidicula cymbalum, arrows point to the outer rim (left) and the inner rim.
Old test, with relatively broad rim, 89 µm – Heelsumse beek, the Netherlands
Pyxidicula cymbalum
Detail of a 95 µm large shell
Pyxidicula cymbalum
Pyxidicula cymbalum
Pyxidicula cymbalum
Pyxidicula cymbalum,c. 75 µm
Pyxidicula cymbalum
Pyxidicula cymbalum
Pyxidicula cymbalum c. 69 µm
Pyxidicula cymbalum
Pyxidicula cymbalum
Pyxidicula cymbalum
Pyxidicula cymbalum c. 82 µm
Pyxidicula cymbalum
Pyxidicula cymbalum, diam. 119 µm, Lilla Kroksjön, Sweden, 2026
Ferry Siemensma, created February 27, 2019; last modified May 17, 2026
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