
Clathrella foreli, with nucleus visible, Crailoo, Netherlands, 2024
Clathrella foreli Penard, 1903
Diagnosis: Cell spherical, surrounded by numerous hyaline, colorless, bowl‑shaped scales that are loosely attached to one another. Filopodia emerge through the gaps between the scales. Pseudopodia are branching and, when attached to the substrate, may also anastomose. One vesicular nucleus.
Dimensions: Penard: 40–55 µm (adult amoebae). Hoogenraad & De Groot (1940): 40–50 µm. My measurements: 25–86 µm, mean 35 µm (body only, without scales). Scales 9.8–16 µm in diameter; nucleus 8.2–16 µm.
Ecology: Oligotrophic and mesotrophic freshwater habitats. Recorded from Europe, North and South America, Japan, China, and India.
Remarks: For additional information, see my paper in Mikrokosmos (Siemensma, 2014)

Clathrella foreli, Crailoo, Netherlands, 2024

Clathrella foreli, Crailoo, Netherlands, 2024

Surface structure (Spiegelplas, Netherlands, 2023)


Two specimens from Crailoo, a mesotrophic freshwater pond

Clathrella foreli, n=nucleus, plasmabody 30 µm, scales about 14 µm – Peru, rainforest lagoon.

Clathrella foreli, 31 µm, nucleus clearly visible.

Specimen showing the polygonal structure of the scales and the branching and anastomosing filopodia.

The arrow points to a gap between the scales through which pseudopodia pass.

Specimen with many branching pseudopodia



Clathrella foreli, 30 µm – Peru, feeding on diatoms


Empty shell


Clathrella foreli, dried shell, from Naardermeer and Peru


Clathrella foreli, 32 µm, Naardermeer


Clathrella foreli, Naardermeer and Crailoo


Clathrella foreli, body 40 and 32 µm, Montana USA

Clathrella foreli,sediment in a lake at the Great Alvar, Öland, Sweden, 2026